DOI
https://dx.doi.org/10.18590/euscorpius.2015.vol2015.iss203.1
Abstract
The taxonomic status of the Anatolian-Caucasian subspecies of Euscorpius mingrelicus (Kessler, 1874) is reconsidered. We restrict E. mingrelicus s.str. to Georgia and Turkey, and elevate to species status three of its subspecies: E. phrygius Bonacina, 1980, stat. n., E. uludagensis Lacroix, 1995, stat. n., and E. ciliciensis Birula, 1898, stat. n. We synonymize E. mingrelicus legrandi Lacroix, 1995 with E. phrygius stat. n., and E. m. ollivieri Lacroix, 1995, with E. mingrelicus.
References with DOI
BIRULA, A. A. 1896. Miscellanea scorpiologica. I. Zur Synonymie der russischen Skorpione. Annuaire du Musée Zoologique de l´Académie Impériale des Sciences de St.-Pétersbourg, 1: 229–245.
BIRULA, A. A. 1898. Ein Beitrag zur Kenntniss der Skorpionen fauna Kleinasiens. Horae Societatis Entomologicae Rossicae, 33: 132–140.
BIRULA, A. A. 1900. Scorpiones mediterranei Musei Zoologici mosquensis. Izvestiya Imperatorskogo Obshchestva Lyubitelei Prirody, Istorii, Antropologii i Etnografii (Societas Caesarea Amicorum Rerum Naturalium, Anthropologiae Ethnographiae Universitatis Moscoviensis), 98, 3(1): 8–20 (in Russian).
BIRULA, A. A. [BYALYNITSKII-BIRULYA, A. A.] 1917a. Arachnoidea Arthrogastra Caucasica. Pars 1. Scorpiones. Zapiski Kavkazskago Muzeya [Mémoires du Musée du Caucase]. Series A, No. 5. Tiflis: Imprimerie de la Chancellerie du Comité du Transcaucasie. 253 pp. (in Russian; published August 1917). English translation: ByalynitskiiBirulya, A. A. 1964. Arthrogastric Arachnids of Caucasia. Part 1. Scorpions. Tiflis: Annals of the Caucasian Museum. Series A, No. 5. Jerusalem: Israel Program for Scientific Translations, 170 pp.
BIRULA, A. A. [BYALYNITSKII-BIRULYA, A. A.] 1917b. Faune de la Russie et des pays limitrophes fondee principalement sur les collections du Musée Zoologique de l'Académie des Sciences de Russie. Arachnides (Arachnoidea). Petrograd, 1(1): xx, 227 pp. (in Russian; Introduction dated October 1917). English translation: Byalynitskii-Birulya, A. A. 1965. Fauna of Russia and Adjacent Countries. Arachnoidea. Vol. I. Scorpions. Jerusalem: Israel Program for Scientific Translations, xix, 154 pp.
BONACINA, A. 1980. Sistematica specifica e sottospecifica del complesso “Euscorpius germanus” (Scorpiones, Chactidae). Rivista del Museo Civico di Scienze Naturali "Enrico Caffi" (Bergamo) 2: 47–100.
CRUCITTI, P. 1999. The scorpions of Anatolia: biogeographical patterns. Biogeographia, 20: 81–94.
CRUCITTI, P. & D. CICUZZA. 2001. Scorpions of Anatolia: ecological patterns. Pp. 225–234 In: Fet, V. & Selden, P.A. (eds). Scorpions 2001. In memoriam Gary A. Polis. British Arachnological Society, Burnham Beeches, Bucks.
DI CAPORIACCO, L. 1950. Le specie e sottospecie del genere “Euscorpius” viventi in Italia ed in alcune zone confinanti. Memorie /Atti della Accademia Nazionale dei Lincei, serie VIII, vol. II, sez. III, fasc. 4: 159–230.
FET, V. 1985. Notes on some Euscorpius (Scorpions: Chactidae) from Greece and Turkey. Rivista del Museo Civico di Scienze Naturali “Enrico Caffi” (Bergamo), 9: 3–11.
FET, V. 1989. A catalogue of scorpions (Chelicerata: Scorpiones) of the USSR. Rivista del Museo Civico di Scienze Naturali “Enrico Caffi” (Bergamo), 13: 73–171.
FET, V. 1993. Notes on Euscorpius mingrelicus (Kessler, 1874) from the Caucasus. Rivista del Museo Civico di Scienze Naturali "Enrico Caffi" (Vergamo), 16: 1–8.
FET, V. 2000. Scorpions (Arachnida, Scorpiones) from the Balkan Peninsula in the collections of the National Museum of Natural History, Sofia. Historia Naturalis Bulgarica, 11: 47–60.
FET, V. 2010. Scorpions of Europe. Acta Zoologica Bulgarica, 62(1): 3–12.
FET, V., AY. KARATAŞ, E. V. FET & A. KARATAŞ. 2003. First data on the molecular phylogeny of Euscorpius (Scorpiones: Euscorpiidae) from Turkey. Zoologicheskii Zhurnal, 82(12): 1518–1521 (Moscow, Russia) (in Russian; English summary) (English translation: Entomological Review, 2003, 83, Suppl. 2: 249-252).
FET, V. & W. D. SISSOM. 2000. Family Euscorpiidae Laurie, 1896. Pp. 355–380 in: Fet, V., W. D. Sissom, G. Lowe & M. E. Braunwalder (eds.). Catalog of the Scorpions of the World (1758-1998). New York: The New York Entomological Society.
FET, V. & M. E. SOLEGLAD, 2007. Fauna and zoogeography of scorpions (Arachnida: Scorpiones) in Bulgaria. Pp. 405–422 In: Fet, V. & A. Popov (eds). Biogeography and Ecology of Bulgaria. Springer.
FET, V., M. E. SOLEGLAD, B. GANTENBEIN. 2004. The Euroscorpion: genus Euscorpius (Scorpiones: Euscorpiidae). Proceedings of the 3d Scorpiology Symposium, American Arachnological Society 28th Annual Meeting, Norman, Oklahoma, 23–27 June 2004. Euscorpius, 17: 47–59. https://doi.org/10.18590/euscorpius.2004.vol2004.iss17.1
FET, V., M. E. SOLEGLAD, A. PARMAKELIS, P. KOTSAKIOZI & I. STATHI. 2014. Two new species of Euscorpius from Euboea Island, Greece (Scorpiones: Euscorpiidae). Arthropoda Selecta, 23: 11–26. https://doi.org/10.15298/arthsel.23.2.03
GANTENBEIN, B., V. FET, C. R. LARGIADÈR & A. SCHOLL. 1999. First DNA phylogeny of Euscorpius Thorell, 1876 (Scorpiones: Euscorpiidae) and its bearing on taxonomy and biogeography of this genus. Biogeographica (Paris), 75: 49–65.
HJELLE, J. T. 1990. Anatomy and morphology. Pp. 9– 63 in: Polis, G.A. (ed.), Biology of Scorpions. Stanford, CA: Stanford University Press.
KALTSAS, D., I. STATHI & V. FET. 2008. Scorpions of the Eastern Mediterranean. Pp. 209-246 in Makarov S.E, Dimitrijević R.N. (eds). Advances in Arachnology and Developmental Biology. Papers dedicated to Prof. Dr. Božidar Ćurčić. Vienna– Belgrade–Sofia, 517 pp.
KAYA, U., E. ÇEVIK & U. C. ERIŞMIŞ. 2005. Population status of the Taurus frog, Rana holtzi Werner (1898), in its terra typica: is there a decline? Turkish Journal of Zoology, 29: 317–319.
KESSLER, K. F. 1874. On Russian scorpions, Trudy Russkago Entomologicheskago Obshchestva v S.- Peterburge (Transactions of the Russian Entomological Society in St. Petersburg), 8: 3–27. (in Russian).
KINZELBACH, R. 1975. Die Skorpione der Ägäis. Beiträge zur Systematik, Phylogenie und Biogeographie. Zoologische Jahrbücher, Abteilung für Systematik, 102: 12–50.
KINZELBACH, R. 1982. Die Skorpionssammlung des Naturhistorischen Museums der Stadt Mainz. Teil I: Europa und Anatolien. Mainzer naturwisserschaftliches Archiv, 20: 49–66.
KINZELBACH, R. 1985. Vorderer Orient. Scorpione (Arachnida: Scorpiones). Tübinger Atlas der Vorderer Orients (TAVO), Karte Nr. A VI 14.2.
KOVAŘÍK, F., V. FET, M. E. SOLEGLAD & E. A. YAĞMUR. 2010. Etudes on iurids, III. Revision of the genus Iurus Thorell, 1876 (Scorpiones: Iuridae), with a description of two new species from Turkey. Euscorpius, 95: 1–212. https://doi.org/10.18590/euscorpius.2010.vol2010.iss95.1
KRITSCHER, E. 1993. Ein Beitrag zur Verbreitung der Skorpione im Östlichen Mittelmeerraum. Annalen des Naturhistorischen Museums in Wien. Serie B. Botanik und Zoologie, 94/95, B: 377–391.
KULCZYŃSKI, W. 1903. Arachnoidea in Asia Minore et ad Constantinopolim a Dre. F. Werner collecta. Sitzungsberichte der K. Akademie der Wissenschaften, Mathematisch-naturwissenschaftliche Klasse, Vienna, 112, part I (7): 627–680, 1 unnumbered plate.
LACROIX, J.-B. 1991. Faune de France; Arachnida: Scorpionida. 5e note. Sub-genus (Euscorpius) Thorell, 1876. Arachnides, 8: 17–36.
LACROIX, J.-B. 1995. Euscorpius (E.) mingrelicus Kessler, 1876 en Turquie anatolienne (Arachnida: Scorpionida). Arachnides, 26: 4–6.
PARMAKELIS, A., P. KOTSAKIOZI, I. STATHI, S. POULIKARAKOU & V. FET. 2013. Hidden diversity of Euscorpius (Scorpiones: Euscorpiidae) in Greece revealed by multilocus species-delimitation approaches. Biological Journal of the Linnean Society, 110: 728–748. https://doi.org/10.1111/bij.12170
SCHERABON, B., B. GANTENBEIN, V. FET, M. BARKER, M. KUNTNER, C. KROPF & D. HUBER. 2000. A new species of scorpion from Austria, Italy, Slovenia and Croatia: Euscorpius gamma Caporiacco, 1950, stat. nov. (Scorpiones: Euscorpiidae). In: Gajdoš P. & Pekár S. (eds). Proceedings of the 18th European Colloquium of Arachnology, Stará Lesná, 1999. Ekológia (Bratislava), 19: 253–262. SISSOM, W. D. 1990. Systematics, biogeography and paleontology. Pp. 64–160 in: Polis G. A. (ed.) Biology of Scorpions. Stanford, CA: Stanford University Press.
SOLEGLAD, M. E. & V. FET. 2003. The scorpion sternum: structure and phylogeny (Scorpiones: Orthosterni). Euscorpius, 5: 1–33. https://doi.org/10.18590/euscorpius.2003.vol2003.iss5.1
SOLEGLAD, M. E. & W. D. SISSOM. 2001. Phylogeny of the family Euscorpiidae Laurie, 1896: a major revision. Pp. 25–112 in: Fet, V. & P. A. Selden (eds.) Scorpions 2001. In Memoriam Gary A. Polis. Burnham Beeches, Bucks, UK: British Arachnological Society.
STAHNKE, H. L. 1971. Scorpion nomenclature and mensuration. Entomological News, 81: 297–316. TARKOWSKI, R. 2011. Konstanty Jelski (1837–1896). Przyrodnik i badacz Ameryki Południowej. Kraków: Wydawnictwo Naukowe Uniwersytetu Pedagogicznego, 256 pp.
TROPEA, G. 2013. Reconsideration of the taxonomy of Euscorpius tergestinus (Scorpiones: Euscorpiidae). Euscorpius, 162: 1−23. https://doi.org/10.18590/euscorpius.2013.vol2013.iss162.1
TROPEA, G. & V. FET. 2015. Two new Euscorpius species from central-western Greece (Scorpiones: Euscorpiidae). Euscorpius, 199: 1–16. https://doi.org/10.18590/euscorpius.2015.vol2015.iss199.1
TROPEA, G., V. FET, A. PARMAKELIS, P. KOTSAKIOZI & I. STATHI. 2015. A new species of Euscorpius (Scorpiones: Euscorpiidae) from southern Bulgaria. Arachnologische Mitteilungen, 49: 10–20. https://doi.org/10.5431/aramit4902
TROPEA, G., E. A. YAĞMUR & F. YEŞILYURT. 2014. A new species of Euscorpius Thorell, 1876 (Scorpiones, Euscorpiidae) from the Antalya Province, Southern Turkey. Euscorpius, 184: 1–13. https://doi.org/10.18590/euscorpius.2014.vol2014.iss184.1
VACHON, M. 1974. Etude des caractères utilisés pour classer les familles et les genres de scorpions (Arachnides). 1. La trichobothriotaxie en arachnologie. Sigles trichobothriaux et types de trichobothriotaxie chez les scorpions. Bulletin du Muséum national d’Histoire naturelle, Paris, 140: 859–958.
VACHON, M. & R. KINZELBACH. 1987. On the taxonomy and distribution of the scorpions of the Middle East, In: Krupp, F., W. Schneider & R. Kinzelbach (Eds.). Proceedings of the Symposium on the Fauna and Zoogeography of the Middle East, Mainz (TAVO), 28: 91–103.
VIGNOLI, V. & N. SALOMONE. 2008. A review of and additions to the current knowledge of the scorpion genus Euscorpius Thorell, 1876. Fragmenta Entomologica, 40: 189–228. https://doi.org/10.4081/fe.2008.93
WERNER, F. 1898. Ueber einige neue Reptilien und einen neuen Frosch aus dem Cilicischen Taurus. Zoologischer Anzeiger, 21: 216–223.
WERNER, F. 1902. Die Skorpione, Pedipalpen und Solifugen in der zoologisch-vergleichend-anatomischen Sammlung der Universität Wien. Verhandlungen der Zoologisch-Botanischen Gesellschaft in Wien, 52: 595–608.
Recommended Citation
Tropea, G., Yağmur, E. A., & Fet, V. 2015 . A revision of the Anatolian-Caucasian “Euscorpius mingrelicus complex” (Scorpiones: Euscorpiidae). Euscorpius, No. 203: 1-32.
