A protein fold switch joins the circadian oscillator to clock output in cyanobacteria

Document Type

Article

Publication Date

7-2015

Abstract

Organisms are adapted to the relentless cycles of day and night, because they evolved timekeeping systems called circadian clocks, which regulate biological activities with ~24-hour rhythms. The clock of cyanobacteria is driven by a three-protein oscillator composed of KaiA, KaiB, and KaiC, which together generate a circadian rhythm of KaiC phosphorylation. We show that KaiB flips between two distinct three-dimensional folds, and its rare transition to an active state provides a time delay that is required to match the timing of the oscillator to that of Earth’s rotation. Once KaiB switches folds, it binds phosphorylated KaiC and captures KaiA, which initiates a phase transition of the circadian cycle, and it regulates components of the clock-output pathway, which provides the link that joins the timekeeping and signaling functions of the oscillator.

Comments

The version of record is available from the publisher at https://www.science.org/doi/epdf/10.1126/science.1260031.

Copyright © 2015, American Association for the Advancement of Science. All rights reserved.

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