The high-level systematics and phylogeny of the scorpion superfamily Buthoidea are unresolved. A new, formerly undetected character is reported here: the relative alignment of d3 trichobothrium and DMc carina on the pedipalp patella. This and other characters are discussed in the context of possible high-level divisions of Buthoidea. Based on this analysis, we suggest a breakdown of Buthoidea into six proposed tentative phylogenetic groups: Buthus group, Ananteris group, Isometrus group, Charmus group, Uroplectes group, and Tityus group.
Fet, V., Michael, S. E., & Lowe, G. 2005 . A new trichobothrial character for the high-level systematics of Buthoidea (Scorpiones: Buthida). Euscorpius, No. 23: 1-40.