DOI
http://dx.doi.org/10.18590/euscorpius.2016.vol2016.iss236.1
Abstract
Northwestern African Buthacus species are revised. Buthacus stockmanni sp. n. from Morocco and Western Sahara is described and fully complemented with color photos of live and preserved specimens, as well as their habitat. The hemispermatophore of B. stockmanni sp. n. is illustrated and described. In addition to morphology and hemi-spermatophores, we also describe the karyotypes of B. stockmanni sp. n. (2n=20). B. stockmanni sp. n. is compared with B. occidentalis Vachon, 1953 and B. ziegleri Lourenço, 2000. Both these species are differentiated from B. stockmanni sp. n. geographically and morphologically. The male of B. stockmanni sp. n. has fingers of pedipalp chela strongly twisted proximally and males of the other two species have fingers straight or almost straight. Buthacus huberi Lourenço, 2001 is synonymized with Buthacus occidentalis Vachon, 1953; Buthacus mahraouii Lourenço, 2004 and Buthacus leptochelys algerianus Lourenço, 2006 are synonymized with Buthacus ziegleri Lourenço, 2000.
References with DOI
BIRULA, A.A. 1908. Ergebnisse der mit Subvention aus der Erbschaft Treitl unternommenen zoologischen Forschungsreise Dr. F. Werner`s nach dem ägyptischen Sudan und Nord-Uganda. XIV. Scorpiones und Solifugae. Sitzungsberichte der KaiserlichKöniglichen Akademie der Wissenschaften, Wien, 117/2 (1): 121–152.
FET, V. & G. LOWE. 2000. Family Buthidae C. L. Koch, 1837. Pp. 54–286 in Fet, V., W. D. Sissom, G. Lowe & M. E. Braunwalder. Catalog of the Scorpions of the World (1758–1998). New York: The New York Entomological Society, 689 pp.
FET, V., M.E. SOLEGLAD & G. LOWE. 2005. A new trichobothrial character for the high–level systematics of Buthoidea (Scorpiones: Buthida). Euscorpius, 23: 1–40. https://doi.org/10.18590/euscorpius.2005.vol2005.iss23.1
HEMPRICH, F.W. & C.G. EHRENBERG. 1828. Zoologica II. Arachnoidea. In: Symbolae Physicae seu Icones et Descriptiones Animalium Evertebratorum Sepositis Insectis quae ex Itinere per Africam Borealem et Asiam Occidentalem. Friderici Guilelmi Hemprich et Christiani Godofredi Ehrenberg, Medicinae et Chirurgiae Doctorum, Studio Novae aut Illustratae Redierunt. Percensuit et Regis Iussu et Impensis Edidit Dr. C.G. Ehrenberg. Decas Prima. Berolini ex Officina Academica, Venditur a Mittlero: plate I Buthus; plate II: Androctonus.
HEMPRICH, F. W. & C.G. EHRENBERG. 1829. Vorläufige Uebersicht der in Nord-Afrika und WestAsien einheimischen Skorpione und deren geographischen Verbreitung. Verhandungen der Gesellschaft der Naturforschender Freunde in Berlin, 1(6): 348–362.
KOVAŘÍK, F. 2005. Taxonomic position of species of the genus Buthacus Birula, 1908 described by Ehrenberg and Lourenço, and description of a new species (Scorpiones: Buthidae). Euscorpius, 28: 1– 13. https://doi.org/10.18590/euscorpius.2005.vol2005.iss28.1
KOVAŘÍK, F. 2009. Illustrated catalog of scorpions. Part I. Introductory remarks; keys to families and genera; subfamily Scorpioninae with keys to Heterometrus and Pandinus species. Prague: Clairon Production, 170 pp.
KOVAŘÍK, F., G. LOWE, J. PLÍŠKOVÁ & F. ŠŤÁHLAVSKÝ. 2013. A new scorpion genus, Gint gen. n., from the Horn of Africa (Scorpiones, Buthidae). Euscorpius, 173: 1–19. https://doi.org/10.18590/euscorpius.2013.vol2013.iss173.1
KOVAŘÍK, F., G. LOWE, K. B. RANAWANA, D. HOFEREK, V. A. SANJEEWA JAYARATHNE, J. PLÍŠKOVÁ & F. ŠŤÁHLAVSKÝ. 2016. Scorpions of Sri Lanka (Arachnida, Scorpiones: Buthidae, Chaerilidae, Scorpionidae) with description of four new species of the genera Charmus Karsch, 1879 and Reddyanus Vachon, 1972 stat. n.. Euscorpius, 220: 1–133. https://doi.org/10.18590/euscorpius.2016.vol2016.iss220.1
KOVAŘÍK, F & A.A. OJANGUREN AFFILASTRO. 2013. Illustrated catalog of scorpions Part II. Bothriuridae; Chaerilidae; Buthidae I., genera Compsobuthus, Hottentotta, Isometrus, Lychas, and Sassanidotus. Prague: Clairon Production, 400 pp.
KOVAŘÍK, F., F. ŠŤÁHLAVSKÝ, T. KOŘÍNKOVÁ & J. KRÁL. 2009. Tityus ythieri Lourenço, 2007 is a synonym of Tityus magnimanus Pocock, 1897 (Scorpiones: Buthidae): a combined approach using morphology, hybridization experiments, chromosomes, and mitochondrial DNA. Euscorpius, 77: 1– 12. https://doi.org/10.18590/euscorpius.2009.vol2009.iss77.1
KRAEPELIN, K. 1891. Revision der Skorpione. I. Die Familie des Androctonidae. Jahrbuch der Hamburgischen Wissenschaftlichen Anstalten, 8(1890): 144–286 (1–144).
LEVY, G., P. AMITAI & A. SHULOV. 1973. New scorpions from Israel, Jordan and Arabia. Zoological Journal of the Linnaean Society, 52: 113– 140. https://doi.org/10.1111/j.1096-3642.1973.tb00782.x
LOURENÇO, W.R. 2000. A new species of Buthacus Birula from Morocco (Arachnida: Scorpiones: Buthidae). Faunistische Abhandlungen Staatliches Museum für Tierkunde Dresden, 22(1): 5–9.
LOURENÇO, W.R. 2001. Further taxonomic considerations on the Northwestern African species of Buthacus Birula (Scorpiones, Buthidae), and description of two new species. Entomologische Mitteilungen aus dem Zoologischen Museum Hamburg, 13(163): 255–269.
LOURENÇO, W.R. 2004. New considerations on the Northwestern African species of Buthacus Birula (Scorpiones: Buthidae), and description of a new species. Revista Ibérica de Aracnología, 10: 225– 231.
LOURENÇO, W.R. 2006. Further considerations on the genus Buthacus Birula, 1908 (Scorpiones: Buthidae), with a description of one new species and two new subspecies. Boletín de la Sociedad Entomológica Aragonesa, 38: 59–70.
LOURENÇO W.R. 2013. The Buthacus Birula, 1908 populations from Tassili n'Ajjer (Scorpiones, Buthidae) and description of a new species. Entomologische Mitteilungen aus dem Zoologischen Museum Hamburg, 16 (190) : 89–99.
LOURENÇO, W.R. & E.-A. LEGUIN. 2009. A new species of the genus Buthacus Birula, 1908 from the United Arab Emirates (Scorpiones, Buthidae). Zoology in the Middle East, 46: 103–108. https://doi.org/10.1080/09397140.2009.10638335
LOURENÇO, W.R. & J.-X. QI. 2007. A new species of Buthacus Birula, 1908 (Scorpiones, Buthidae) from Mali. Entomologische Mitteilungen aus dem Zoologischen Museum Hamburg, 14(176): 387–392.
MATTOS, V.F., D.M. CELLA, L.S. CARVALHO, D.M. CANDIDO & M.C. SCHNEIDER. 2013. High chromosome variability and the presence of multivalent associations in buthid scorpions. Chromosome Research, 21: 121–136. https://doi.org/10.1007/s10577-013-9342-3
SIMON, E. 1910. Révision des scorpions d´Egypte. Bulletin de la Société Entomologique d´Égypte, 1910: 57–87. STAHNKE, H.L. 1971. Scorpion nomenclature and mensuration. Entomological News, 81: 297–316.
STOCKMANN, M., C. TURIEL, F. ALTHOFF, G. LOWE & F. KOVAŘÍK. 2016. First Report of Lissothus occidentalis Vachon, 1950 (Scorpiones: Buthidae) from Morocco and Western Sahara, with Notes on Ecology and Captive Breeding. Euscorpius, 235: 1–12. https://doi.org/10.18590/euscorpius.2016.vol2016.iss235.1
VACHON, M. 1953. Contribution á l`étude du peuplement de la Mauritanie. Scorpions. Bulletin d´Institut Français d´Afrique Noire, 55(3): 1012– 1028.
VACHON, M. 1974. Études des caractères utilisés pour classer les familles et les genres des scorpions (Arachnides). 1. La trichobothriotaxie en arachnologie. Sigles trichobothriaux et types de trichobothriotaxie chez les Scorpions. Bulletin du Muséum national d’Histoire naturelle, 3e série, 140 (Zoologie, 104): 857–958.
ZAMBRE, A.M. & W.R. LOURENÇO. 2010. A new species of Buthacus Birula, 1908 (Scorpiones: Buthidae) from India. Boletin Sociedad Entomológica Aragonesa, 46: 115–119.
Recommended Citation
Kovařík, F., Lowe, G., & Šťáhlavský, F. 2016 . Review of Northwestern African Buthacus, with description of Buthacus stockmanni sp. n. from Morocco and Western Sahara (Scorpiones, Buthidae). Euscorpius, No. 236: 1-18.
