DOI
https://dx.doi.org/10.18590/euscorpius.2019.vol2019.iss288.1
Abstract
The diagnostic characters are reassessed and defined for the genera Lychas C. L. Koch, 1845, Mesobuthus Vachon, 1950, and Olivierus Farzanpay, 1987 (the latter is restored from synonymy with Mesobuthus). Four new genera are described: Aegaeobuthus gen. n. (type species Buthus gibbosus Brullé, 1832), Afrolychas gen. n. (type species Isometrus burdoi Simon, 1882), Janalychas gen. n. (type species Lychas srilankensis Lourenço, 1997), and Spelaeolychas gen. n. (type species Isometrus hosei Pocock, 1891). Type species are designated for subgenera Lychas (Distotrichus) Tikader & Bastawade, 1983 (type species Isometrus nigristernis Pocock, 1899), Lychas (Alterotrichus) Tikader & Bastawade, 1983 (type species Scorpio mucronatus Fabricius, 1793), and Lychas (Endotrichus) Tikader & Bastawade, 1983 (type species Isometrus scaber Pocock, 1893). All these three subgenera are now in synonymy with Lychas C. L. Koch, 1845. Lychas kaimana Lourenço, 2011 is synonymized with Lychas shelfordi (Borelli, 1904). Taxonomic position of Lychas timorensis Lourenço, 2018, which is a member of Lychas variatus (Thorell, 1876) complex, is discussed. The species and subspecies of Mesobuthus Vachon, 1950 are discussed, with seven subspecies elevated to species level: Mesobuthus afghanus (Pocock, 1889), stat. n., M. bogdoensis (Birula, 1896), stat. n., M. haarlovi Vachon, 1959, stat. n., M. iranus (Birula, 1917), stat. n., M. mongolicus (Birula, 1912), stat. n., M. persicus (Pocock, 1899), stat. n., and M. thersites (C. L. Koch, 1839), stat. n. Taxonomic changes are supported by 182 figures including the first published photographs of the syntypes of Olivierus hainanensis (Birula, 1904), stat. n., comb. n. and O. przewalskii (Birula, 1897), comb. n.
References with DOI
Farzanpay, R. 1987 (1366). [Knowing Scorpions]. Teheran: Central University Publications, No. 312, Biology 4, 231 pp. (in Farsi, with Latin index).
Fet, V. 1997. Notes on the taxonomy of some old world scorpions (Scorpiones: Buthidae, Chactidae, Ischnuridae, Scorpionidae). The Journal of Arachnology, 25: 245–250.
Fet, V., F. Kovařík, B. GantenbEIn, R. C. KaIser, A. K. Stewart & M. R. Graham. 2018. Revision of the Mesobuthus caucasicus complex from Central Asia, with descriptions of six new species (Scorpiones: Buthidae). Euscorpius, 255: 1–77. https://doi.org/10.18590/euscorpius.2018.vol2018.iss255.1
FET, V. & G. LOWE. 2000. Family Buthidae C. L. Koch, 1837. Pp. 54–286 in Fet, V., W. D. Sissom, G. Lowe & M. E. Braunwalder. Catalog of the Scorpions of the World (1758–1998). New York: The New York Entomological Society, 689 pp.
FRANCKE, O. F. 1985. Conspectus genericus scorpionorum 1758–1982 (Arachnida: Scorpiones). Occasional Papers of the Museum, Texas Tech University, 98: 1–32. https://doi.org/10.5962/bhl.title.142852
Gantenbein, B., V. FET, I. A. Gantenbein-Ritter & F. Balloux. 2005. Evidence for recombination in scorpion mitochondrial DNA (Scorpiones: Buthidae). Proceedings of the Royal Society: Biological Sciences, 272(1564): 697–704. https://doi.org/10.1098/rspb.2004.3017
GANTENBEIN, B., V. FET & A. V. GROMOV. 2003. The first DNA phylogeny of four species of Mesobuthus Vachon, 1950 (Scorpiones: Buthidae) from Eurasia. Journal of Arachnology, 31: 412–420. https://doi.org/10.1636/h01-23
Koch, C. L. 1837. Üebersicht des Arachnidensystems. Erstes Heft. Nürnberg: C. H. Zeh’schen Buchhandlung., 39pp. https://doi.org/10.5962/bhl.title.39561
Koch, C. L. 1845. Die Arachniden. Nürnberg: C. H. Zech’sche Buchhandlung, 11, 174 pp. Koch, C. L. 1850. Übersicht des Arachnidensystems, 5. Nürnberg: C. H. Zeh’schen Buchhandlung., 104 pp.
Koch, L. E. 1977. The taxonomy, geographic distribution and evolutionary radiation of Australo-Papuan scorpions. Records of the Western Australian Museum, 5(2): 83–367.
Kovařík, F. 1995. Review of Scorpionida from Thailand with descriptions of Thaicharmus mahunkai gen. et sp. n. and Lychas krali sp. n. (Buthidae). Acta Societatis Zoologicae Bohemicae, 59: 187–207.
KOVAŘÍK, F. 2009. Illustrated catalog of scorpions. Part I. Introductory remarks; keys to families andgenera; subfamily Scorpioninae with keys to Heterometrus and Pandinus species. Prague: Clairon Production, 170 pp.
Kovařík, F. G. Lowe, K. B. Ranawana, D. Hoferek, V. A. Sanjeewa Jayarathne, J. Plíšková & F. Šťáhlavský 2016. Scorpions of Sri Lanka (Arachnida, Scorpiones: Buthidae, Chaerilidae, Scorpionidae) with description of four new species of the genera Charmus Karsch, 1879 and Reddyanus Vachon, 1972 stat. n. Euscorpius, 220: 1–133. https://doi.org/10.18590/euscorpius.2016.vol2016.iss220.1
KOVAŘÍK, F. & A. A. OJANGUREN AFFILASTRO. 2013. Illustrated catalog of scorpions. Part II. Bothriuridae; Chaerilidae; Buthidae I. Genera Compsobuthus, Hottentotta, Isometrus, Lychas, and Sassanidotus. Prague: Clairon Production, 400 pp.
KOVAŘÍK, F. 2018. Notes on the genera Buthacus, Compsobuthus, and Lanzatus with several synonymies and corrections of published characters (Scorpiones: Buthidae). Euscorpius, 269: 1–12. https://doi.org/10.18590/euscorpius.2018.vol2018.iss269.1
KraepelIN, K. 1891. Revision der Skorpione. I. Die Familie des Androctonidae. Jahrbuch der Hamburgischen Wissenschaftlichen Anstalten, 8(1890): 144–286 (1–144).
Lourenço, W. R. 1997. A new species of Lychas Koch, 1845 (Chelicerata, Scorpiones, Buthidae) from Sri Lanka. Revue suisse de Zoologie, 104(4): 831–836. https://doi.org/10.5962/bhl.part.80024
Lourenço, W. R. 2005. A new genus and species of scorpion from Afghanistan (Scorpiones, Buthidae). Bonner Zoologische Beiträge, 53(2004): 111–114.
LOURENÇO, W. R. 2011. Scorpions from West Papua, Indonesia and description of a new species of Lychas C. L. Koch, 1845 (Scorpiones: Buthidae). Entomologische Mitteilungen aus dem Zoologischen Museum Hamburg, 15(186): 317–326.
LOURENÇO, W. R. 2018. Descriptions of a new species of Lychas C. L. Koch, 1845 (Scorpiones: Buthidae) from East Timor and the male of L. eliseanneae Lourenco, 2011 from West Papua. Arachnida - Rivista Aracnologica Italiana, 19: 27–37.
Mirshamsi, O., A. Sari, E. Elahi & S. Hosseinie. 2011. Mesobuthus eupeus (Scorpiones: Buthidae) from Iran: a polytypic species complex. Zootaxa, 2929: 1–21.
Pocock, R. I. 1891. On some Old-World species of scorpions belonging to the genus Isometrus. Journal of the Linnaean Society, 23: 432–447. https://doi.org/10.1111/j.1096-3642.1891.tb00386.x
Pocock, R. I. 1900. Arachnida. The fauna of British India, including Ceylon and Burma. Published under the authority of the Secretary of State for India in Council. London: W. T. Blandford, xii, 279 pp.
PrendInI, L. 2005. Scorpion diversity and distribution in southern Africa: Pattern and process. Pp. 25–68 in Huber, B.A., B. J. Sinclair & K.-H. Lampe (Eds.) African Biodiversity: Molecules, Organisms, Ecosystems. Proceedings of the 5th International Symposium on Tropical Biology, Museum Alexander Koenig, Bonn. Springer Verlag, New York. https://doi.org/10.1007/0-387-24320-8_2
Purcell, W. F. 1902. On some South African Arachnida belonging to the orders Scorpiones, Pedipalpi, and Solifugae. Annals of South African Museum, 2: 137–225.
SIMON, E. 1882. Arachnides. In L. Fairmaire, L. & E. Simon. Récoltes entomologiques de M. A. Burdo, sur le trajet de Zanzibar aux Grands Lacs. Annales de la Société Entomologique de Belgique, 26: 58–60.
Stahnke, H. L. 1971. Scorpion nomenclature and mensuration. Entomological News, 81(12): 297–316.
Tikader, B. K. & D. B. Bastawade. 1983. Scorpions (Scorpionida: Arachnida). In The Fauna of India, Vol. 3. (Edited by the Director). Calcutta: Zoological Survey of India, 671pp.
Vachon, M. 1952. Études sur les scorpions. Institut Pasteur d’Algérie, Alger, 1–482. (published 1948–1951 in Archives de l’Institut Pasteur d’Algérie, 1948, 26: 25–90, 162–208, 288–316, 441–481. 1949, 27: 66–100, 134–169, 281–288, 334–396. 1950, 28: 152–216, 383– 413. 1951, 29: 46–104).
VACHON, M. 1974. Etude des caractères utilisés pour classer les familles et les genres de Scorpions (Arachnides). 1. La trichobothriotaxie en Arachnologie. Sigles trichobothriaux et types de trichobothriotaxie chez les scorpions. Bulletin du Muséum National d’Histoire Naturelle, Paris, (3), 140 ( Zool. 104), mai-juin 1973: 857–958.
Vachon, M. 1986. Étude de la denture des doigts des pédipalpes chez les scorpions du genre Lychas. Bulletin du Muséum National d’Histoire Naturelle Paris, 8: 835– 850.
Vachon, M. & W. R. LourenÇo. 1985. Scorpions cavernicoles du Sarawak (Borneo). Chaerilus chapmani n. sp. (Chaerilidae) et Lychas hosei (Pocock, 1890) (Buthidae). Mémoires Biospéologiques, 12: 9–18.
YTHIER, E. 2018. A new species of Mesobuthus Vachon, 1950 (Scorpiones: Buthidae) from Crete (Greece). Revista Ibérica de Aracnologia, 32: 87–92.
Recommended Citation
Kovařík, F. 2019 . Taxonomic reassessment of the genera Lychas, Mesobuthus, and Olivierus, with descriptions of four new genera (Scorpiones: Buthidae). Euscorpius, No. 288: 1-27.